Usagi Tsukino is a 14-year-old schoolgirl who is very cheerful, but clumsy and is always finding herself into unwanted trouble. One day, Usagi meets a talking black cat named Luna, who reveals that dark forces are threatening the lives of the innocent and that Usagi has the power to stop them. Using a magical brooch, Usagi is transformed into Sailor Moon, the chosen guardian who fights for love and justice.

A new house of fortune opens up in the Juban District and quickly gains popularity for its accurate predictions. But soon, the customers who had their fortunes read begin to act violently including Gurio Umino, a boy who has a crush on Usagi. Suspecting that this is the work of a monster, Usagi and Luna decide to investigate the situation.

All the girls in town are tuning in to the late-night radio program called "Midnight Zero" that reads out love letters sent in by the listeners. Meanwhile, a strange sleeping sickness is plaguing the town, causing people to fall into a coma-like state, including Ms. Haruna and Naru.

Usagi is shocked to find out that she's gained weight from too much eating. Determined to get back into shape and fast, she and her friends join a gym that advertises instant results for its members.

Usagi asks her family if they can officially keep Luna as a pet, but her little brother Shingo strongly objects due to his fear of cats. Instead, he brings home a pet Chanela, a creature that smells like perfume. Shingo becomes so absorbed in his new pet, that he behaves differently, which worries Usagi.

One rainy day, Usagi bumps into Yusuke Amade, a famous musician who claims he is being chased by a monster. Realizing the man is telling the truth, Usagi and Luna decide to keep an eye on Yusuke by staking out the venue where he performs.

Usagi and Naru both dream of becoming stars like Mikan Shiritori, a graduate from their school. When the girls hear that Mikan is holding a talent contest, they see their chance to achieve stardom.

Usagi befriends Ami Mizuno, the school's smartest girl, who has an IQ of over 300. But Luna, who senses an unknown energy coming from Ami, suspects her of being a monster.

Usagi and Ami go to the city to enjoy some shopping, where they find a clock store selling a clock that resembles Luna. Later that day, Usagi is happy to learn that her mother bought the exact same clock as a gift. But that night, the clocks from the store emit a strange light, making Usagi and the rest of Tokyo very impatient.

Several people who take the bus near the Hikawa Shrine have gone missing. Usagi, Ami, and Luna decide to investigate by visiting the shrine. There, they meet Rei Hino, a shrine maiden with strong spiritual powers.

Having no information on the whereabouts of the princess, the sailor soldiers investigate a new amusement park, which has had a rash of disappearances. While investigating, Rei and Usagi bump into Mamoru. Rei and Mamoru have many things to agree upon because of their "hatred" for Usagi. Ami enters the candy house in Dreamland, the amusement park. The candy house is the reason for the numerous disappearances; it is being used to steal the energy from people.

Jadeite is approached by Tetis with an effective way to collect energy, Operation Romantic Cruise. Using her psychic abilities, Rei wins two tickets to go on the cruise and invites Ami to find boyfriends for themselves. Usagi, very jealous and angry, sneaks onto the ship disguised as a beautiful camera woman. Ami and Rei come to the realization that they are the only single girls on the ship, but can catch guys on the rebound. Jadeite, disguised as the captain of the ship, discovers Usagi, but neither realize who the other is.

Queen Beryl gives Jadeite one last chance to destroy the sailor soldiers. As Usagi gazes into the stars, Jadeite appears in the sky and threatens to burn Tokyo to the ground if the sailor soldiers don't show up at the airport to fight him. The girls decide whether or not to go because they know the airport will be heavily guarded with police. At the airport, Jadeite sees the girls transform and finally knows who they are. Arriving just in time, Tuxedo Mask fights Jadeite and they both fall into the ocean. The girls fear the worst when only Jadeite emerges.

With Jadeite gone, Nephrite explains to Queen Beryl about how he plans to collect energy; he plans to collect energy from individuals at points in their life when their energy is at its peak. Rui is Nephrite's first target. Disguising himself as a tennis coach, Nephrite gives Rui a racket that fills her with negative energy and makes her become obsessed with tennis. Usagi is prompted to investigate after Naru comes to her with concerns about Rui. Just as Usagi approaches, the monster inside Rui's cursed racket is released by Nephrite to collect her energy. Usagi transforms into Sailor Moon to fight the monster. However, she is turned into a giant tennis ball and is bounced around the court!

After discovering a local park is to be demolished, Rei becomes determined to bring a boyfriend there.Usagi becomes upset with Mamoru after he saves Luna's life. Rei decides she is going to go on a date with Mamoru, while Nephrite targets Mr. Kunitachi, the caretaker of the park, as his next victim. Usagi witnesses Rei falling in front of Mamoru, who accidentally steps on her head, and invites him for tea as an apology.

Usagi and Naru discover from Umino that their home economics teacher, Ms. Akiyama, is engaged. Unfortunately, this teacher is Nephrite's newest target to collect energy. Ms. Akiyama decides to enter a handmade wedding dress contest in order to win a grand wedding reception. Usagi and Rei also enter. Rei decides that she can learn from Usagi's mother, a full-time housewife, how to sew, but Ikuko doesn't know how, either! Meanwhile, Nephrite puts a curse on the silk material Ms. Akiyama is going to make her dress out of for the contest.

Girls at Usagi's school are amazed by the pictures taken by a junior high school photographer. Once Kijin Shinokawa, the young photographer, has his camera cursed by Nephrite, he puts out a call for female models. Usagi becomes determined to be one of these models, and talks to Ami, Shingo, Ikuko, and Mamoru about her modeling; she is discouraged by everyone! Kijin lures the unsuspecting models into his studio and "captures" his victims on film. Ignoring Luna's warning, Usagi participates in the contest (even though her suit is full of holes) after she is accepted. Usagi transforms and the monster inside Kijin's camera is released.

Shingo's friend, Mika, is an award-winning doll maker, like her mother. Nephrite targets Mika after winning a French contest. After school, Usagi and Naru are approached by two girls from Shingo's school; they say that he is mean because he broke Mika's award-winning doll without apologizing! Mika has been visited by Nehprite and is under his control. Shingo is saddened by her cool attitude and begins to make a [Sailor Moon] doll of his own to give her as an apology. Later, Usagi, Shingo, Ami, and Rei with Mamoru, go to a huge doll show.

Nephrite writes fake love letters to girls all over the Juuban area in Tuxedo Mask's name, hoping that Sailor Moon will respond. Usagi falls falls for the trick and finds out many other girls have too. Naru decides to meet this mystery man who claims to like her leafing Nephrite to believe Naru is Sailor Moon; he realizes his mistak but proceeds in draining the energy of love Naru produces.

The girls make reservations at a private beach resort to continue their Sailor Warrior training. All they really want to do, however, is have some fun... that is, until they realize that they're booked their stay at the "Adam's Inn". Even though attendants at the Inn look like they're straight out of classic horror movies, the girls make the best of it. Ami tries to befriend Sakiko, the little girl who lives there, but her father rebuffs her. He hypnotizes Sakiko to invoke the powerful spirit inside of her. Unfortunately, the spirit goes out of control and starts attacking everyone.

Nephrite decides to target one of the animators of the Sailor V movies, Hiromi, by cursing one of her pencils. Ami goes to the studio to return a notebook that Hiromi has accidentally dropped, and she sees Nephrite as he is leaving. Suspecting the worst, she brings Usagi and Rei to the studio to investigate. The monster appears and attacks the girls. It is in the form of two twins that are connected ate the tail. They use all of the same techniques as the Sailor Warriors and yet are only defeated when they start bickering about who will give the killing blow.

A visiting princess is holding a ball to show off her nation's treasure, a beautiful jewel. Usagi disguises herself as a princess to invetigate whether this girl could be the missing moon princess. Nephrite also goes because he thinks that the jewel could be the Silver Crystal that Queen Beryl has been seeking. Usagi loves the ball and even gets to dance with Tuxedo Mask (who is also there looking for the crystal). However, Usagi an Tuxedo Mask are left dangling over the side of the balcony when Nephrite possesses the princess. Usagin and Tuxedo float safely to the ground using Usagi's parasol. Later in the evening, Tuxedo Mask visits sailor Moon on the balcony and kisses her under the moonlight. They both remember something like this from the past...

Nephrite gives up trying to get power from humans and focuses on finding the Silver Crystal. He uses a black crystal to help him identify its location. He thinks that Naru (who is still in love with his human guise) may have the clue. However, when he asks for the Silver Crystal, she brings him a jewel from her mother's store. Sailor Moon busts in on the scene just as Nephrite realizes that maybe Naru's love for him is itself a powerful source of energy. A monster sent from Zoisite, who mistakes Nephrites's locating crystal for the Silver Crystal, also appears. Sailor Moon dispatches it and tries to finish Nephrite as well... but Naru throws herself over him, and Sailor Moon is forced to stop her attack.

Nephrite decides that Naru is connected to Sailor Moon after all. When she can't tell him who Sailor Moon really is, he leaves. While he is away, Zoisite's monsters come and kidnap Naru because they think that Nephrite cares for her and will exchange her for the black crystal. Nephrite tracks them down and drives off the monsters, saving Naru. Unfortunately, the monsters return and stick Nephrite with a life-sucking thorn. Naru risks her life to protect Nephrite, but they are both helpless unlit the Sailor Warriors appear.

Queen Beryl wants the Silver Crystal to revive Queen Metalia, the supreme ruler of the Dark Kingdom. Metalia explains that the Silver Crystal is made up of 7 Rainbow Crystals that contain the spirits of 7 powerful monsters. These crystals are sealed away inside of humans. Zoisite uses the black crystal to find the first human carrier. Meanwhile, there is a new girl at school named Makoto. Although she has a bad reputation for starting fights, Usagi makes friends with her because she is an excellent cook. They go to the arcade and see a boy, Joe, win a bunch of prizes.

Usagi, Umino and Naru go on a walk to try and cheer up Naru (who is still upset over Nephrite's death). Naru asks for help from a priest whom they run into at a cemetery. Unfortunately, the priest is the 2nd crystal carrier. Zoisite appears to retrieve the crystal and truns the priest into a monster. Tuxedo Mask arrives and fights Zoisite for the crystal. The other Warriors show up and together they fight the monster and restor his humanity. Sailor Moon notices that Tuxedo Mask left behind a mysterious star shaped locket.

Zoisite discovers that the 3rd crystal carrier is a boy named Ryo Urawa. Ryo is a genius that has just transferred into the Sailor Warriors' school. Ami immediately likes him (a fellow genius!), and they become friends. Urawa reveals to her that he can see the future and that he also knows that Ami is Sailor Mercury and begs her to destroy him before he is turned into a monster. Unfortunately, Zoisite arrives and does just that. The Rainbow Crystal is dropped during the fight, but Sailor Mercury manages to retrieve it.

Usagi and Mamoru are recruited by the artist Yumemi to be models for her next illustration. They both feel drawn to the artists' work because it reminds them of some distant past. Mamoru rebuss her, but Usagi and Umemi become friends. Unfortuanately, Zoisite identifies Umemi as the 4th crystal carrier and attacks her, stealin gthe crystal and turning her into a monster. Tuxedo Mask arrives and protects Sailor Moon, but reveals that he is after the Rainbow Crystals too. He even wants to take away the one that the Sailor Warrios have! Sailor Moon is very confused but somehow manages to defeat the monster and return the artist to normal.

Makoto has a crush on Motoki (the arcade attendant) but is saddened to hear that he already has a girlfriend. Usagi and Makoto get into an argument when Usagi announces that she likes Motoki. Meanwhile, Zoisite targets the girlfriend, Reika, as the 5th crystal carrier. He attacks her and turns her into a monster and steals the crystal. Tuxedo Mask arrives and fights with him for the crystal, but Zoisite tricks him and escapes with it. Sailor Moon and Jupiter work together to fight the monster and restore Reika to normal.

Rei's grandfather is targeted as the 6th crystal carrier. Zoisite appears and tries to extract the crystal, but he is forced to retreat when assaulted by both Grandpa and Rei's two crows. Zoisite's attack, however, awakens the crystal inside of Grandpa and he starts acting weird. Meanwhile, Usagi tries to get Rei and Yuichiro (Grandpa's new apprentice) together as a couple but fails. Zoisite returns to the temple and takes the crystal from Grandpa and turns him into a monster. Yuichiro tries to protect Rei but gets knocked unconscious. Sailor Moon arrives just in time and manages to change Grandpa back without harming him.

Luna is in big trouble when a herd of cats surround and chase her through the city. She is only saved at the last minute by a big blue cat named Red Battler who scares the other cats away. Luna is a bit turned off by the big brute, eve though he obviously has a crush on her. Zoisite realizes that Red Battler is the 7th crystal carrier and attacks. The two cats flee into the sewer but eventually Zoisite catches up with them, captures the crystal and turns the cat into a monster. Tuxedo Mask arrives and managers to wrestle the Rainbow Crystal away from Zoisite. The cat mother still feels something for Luna and doesn't harm her. Sailor Warriors arrive and return the cat to normal.

Umino tries to impress Naru by dressing up as Tuxedo Mask. Zoisite wants to turn Naru into a monster to hopefully draw out Sailor Moon, but is frightened away when he sees Tuxedo Mask (Umino) at her house. He returns the next day, but he misses and turns a nearby performer into a monsters instead. The monster starts to suffocate all of the people (including Umino and Naru), and Zoisite uses this to blackmail Sailor Moon into giving up her Rainbow Crystal. Tuxedo Mask arrives, and they fight for the crystal while Sailor Moon frees the people. In the end, Zoisite grabs it and disappears.

Kunzite and Zoisite work together on a plan to trap Tuxedo Mask and take his crystals. Zoisite disguises himself as Sailor Moon and pretends to be in danger. Tuxedo Mask rescues the fake Sailor Moon and gets stabbed by Zoisite. In the struggle, his mask is torn off and his true identity is revealed to Zoisite. The Sailor Warriors also arrive but get trapped in an energy absorbing dome by Kunzite. Kunzite makes the dome smaller and smaller to force Tuxedo to give up his crystals, but -just in time- Sailor Venus appears with Artemis and rescues the trapped Warriors.

Zoisite tracks down Mamoru and challenges him to a duel where winner takes all of the Rainbow Crystals. Mamoru is heading to duel when he runs into Usagi. She notices his bleeding shoulder and follows him. When they arrive, Zoisite tricks Mamoru into laying down his crystals and Kunzite appears and takes them all. Then Mamoru and Usagi get trapped in the elevator of a crumbling building, and they are both forced to transform (and reveal their identities to each other) in order to save themselves. They find themselves transported to the Dark Kingdom. Zoisite attacks Sailor Moon with a shard, but Tuxedo Mask takes the blow instead. When she sees Tuxedo Mask wounded, Sailor Moon cries. Her tears call all of the Rainbow crystals to her and they form the Silver Crystal. The crystal takes its place on the Moon Stick and Sailor Moon is now transformed into Princess Serenity.

Using the power of the Silver Crystal, Princess Serenity sends Zoisite running. Then, overwhelmed by the memories of her past self, she passes out. With Serenity out, Kunzite appears and takes both Zoisite and the unconscious Tuxedo Mask. When he arrives, Queen Beryl kills Zoisite for his incompetences and turns her attention toward reviving Tuxedo Mask. Back in the cave, Sailor Moon wakes up and finally remembers the past. She and Prince Endymion lived on the moon in a great kingdom called the Silver Millennium. Then a great evil took over the Earth and tried to steal the Silver Crystal. Sailor Moon gets scared and declares that she doesn't want to fight anymore. Then Kunzite returns and blasts the Sailor Warriors. When only Sailor Moon is left, she finally accepts her destiny and calls on the Silver Crystal's power to send Kunzite running.

Usagi is depressed that Mamoru is gone, just when they realized their fate together. Minako tries to cheer her up by taking her to the salon that is offering free hair cuts and styling. The salon is a front for the Darking Kingdom, which is looking for Sailor moons true identity. Minako accidentally gets targeted as Sailor Moon because she has a strand of Usagi's hair on her and is attacked. As Sailor Moon is fighting the monsters she is struck by a black rose thrown by Endymion. Sailor Moon is confused; he appears to be commanding the monsters against her! The other Sailor Warriors appear just in time to drive Tuxedo Mask back and rescue Sailor Moon.

Usagi decides that if she really is the moon princess then she should start acting like one. She and the other girls enroll in a princess training seminar (which is really just a trap for Sailor Moon). Usagi does horribly on all the exercises and fails along with Rei and Minako. After the ""graduates"" are turned into wax statues, the head mistress turns into a monster and attacks the Sailor Warriors under Kunzite and Endymion's supervision. The Sailor Warriors manage to defeat the monster when Kunzite and Endymion start to argue. They escape before Sailor Moon can use the Silver Crystal to heal Endymion's brainwashing.

The girls fall for an advertisement for a ""moon Princess Contest"" and pick up for the mountains. The contest is really a trap for Sailor Moon. The former ""Moon Princess"" Saeko is under the control of the Dark Kingdom and causes and avalanche that traps Rei, Usagi and Yuichiro in the snow. The monster attacks and is stopped at the last minute b a red rose from Endymion (who has momentarily remembered himself). Endymion doesn't stay good for long, however, and soon flees... promising to return and take the Silver Crystal from Sailor Moon.

Kunzite posses two champion ice skaters in an attempt to trap Sailor Moon. He lures the girls to the ice skating rink with free lessons, hoping that the two possesed skaters can identify the moon princess. Makoto skates so well that they think she is Sailor Moon and attack. Sailor Moon transforms and tries to help, but she is still a klutz on the ice. She is saved again by Tuxedo Mask, who disagrees with Kunzite's techniques of manipulating innocents. Sailor Moon and Endymion air up for a skating battle with the monsters. Suddenly, Sailor Moon can skate! Kunzite appears and tires to freeze everyone, but Endymion destroys the freezing machine. Unfortunately, he flees before Sailor Moon can heal him.

Usagi's family goes on vacation to a hot springs resort near the lake where Usagi's parents met. Queen Beryl notices that there is an unusual energy surrounding the lake and sends Endymion to investigate. He discovers that there is a monster buried underneath the lake and uses his powers to revive her. The monster soon starts attacking Usagi's family. Endymion accepts responsibility for unleashing the monster and protects the Tsukinos. When the other Sailor Warriors arrive, he helps them defeat the monster and release the girl's sou. Sadly, he flees again before Sailor Moon can talk to him.